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Blake arrived in St. Louis with his father Gregg Williams, who'd been hired by Jeff Fisher as the Rams defensive coordinator. When the elder Williams fell from grace due to his involvement in the New Orleans Saints "bounty" scandal, the Rams never filled the coordinator void. Instead, Fisher and his staff managed the role by committee. The 27 year old Blake handled defensive play calling duties for the most part, but little is known whether he developed game day defensive schemes.
This will heighten the speculation that Fisher has found his "guy" to fill the defensive coordinator role. Included in the news of Blake's removal is a second hand statement quoting Williams that his father's possible return to the Rams isn't going to happen.